Prepare before arrival
Complete methodological readings and consult with the professor to strengthen the research question, study design, and draft proposal.
Plan a program 
Advanced Primate Behavioral Ecology · Winter Session 2026–2027
An advanced primate field school for students with prior behavioral research experience who are ready to develop and conduct an independent project with wild monkeys under real field conditions.
Course experience
Students prepare through assigned methodological readings, research planning, and faculty consultation before arriving in Costa Rica. At Camaquiri, they refine an approved proposal for research under field conditions and devote most of the course to systematic data collection for an independent project, supported by lectures, group exercises, and regular faculty guidance. During the final week, the group travels to Costa Rica's South Pacific and stays at Casa Gratitud. Students conduct primate censuses, visit Corcovado National Park on the Osa Peninsula, and visit the Indigenous community of Boruca.
The research journey
Students move through the same core stages used in field primatology: careful observation, ecological measurement, an approved research plan, and evidence-based communication.
Complete methodological readings and consult with the professor to strengthen the research question, study design, and draft proposal.
Present the proposal at Camaquiri, evaluate its feasibility on site, and adjust methods when field realities demand it.
Dedicate the core of the field course to systematic behavioral and ecological data collection for an independent project.
Spend the final week at Casa Gratitud, conduct primate censuses, visit Corcovado National Park on the Osa Peninsula, and visit the Indigenous community of Boruca.
Organize and interpret the evidence, connect findings to the scientific literature, and develop the work into a substantial final paper.
Clear from the start
Course tuition is designed to cover the complete academic field experience in Costa Rica during the official program dates.





Meet Your Professor
Dr. Pruetz has directed the Fongoli Savanna Chimpanzee Project in Senegal since 2001. Her comparative research examines how environmental pressures shape ape behavior and inform the study of early hominin ecology.
2026–2027 field courses
Applications begin with fit, preparation, and academic interest. No payment is required to submit the form.
Start your application ↗